1. National Institute of Neurological Disorders and Stroke, National Institutes of Health 1 , Bethesda, MD 20892 , USA
2. Georgetown University 2 Interdisciplinary Program in Neuroscience , , Washington, DC 20007 , USA
3. College of Medicine, University of Kentucky 3 , Lexington, KY 40506 , USA
4. Georgetown University 4 Department of Pathology , , Washington, DC 20007 , USA
5. Georgetown University 5 Department of Neurology , , Washington, DC 20007 , USA