1. Graduate School of Medical and Dental Sciences, Niigata University 1 Division of Neurobiology and Anatomy , , Niigata 951-8510 , Japan
2. Medical Research Institute, Tokyo Medical and Dental University (TMDU) 2 Department of Biochemical Pathophysiology , , Tokyo 113-8510 , Japan
3. National Institute of Neuroscience, National Center of Neurology and Psychiatry (NCNP) 3 Department of Biochemistry and Cellular Biology , , Tokyo 187-8502 , Japan
4. Department of System Pathology for Neurological Disorders, Brain Research Institute, Niigata University 4 , Niigata 9518585 , Japan
5. Center for Coordination of Research Facilities (CCRF), Niigata University 5 , Niigata 951-8510 , Japan