1. Georgetown University 1 , 3700 O St NW, Washington, DC 20097 , USA
2. Carleton University 2 , Ottawa, ON , Canada , K1S 5B6
3. SMRU Consulting, Scottish Oceans Institute 3 , , St Andrews, Fife, KY16 8LB , UK
4. University of St Andrews 3 , , St Andrews, Fife, KY16 8LB , UK
5. Aarhus University 4 Department of Biology , , C. F. Møllers Allé 3, DK-8000 Aarhus C , Denmark
6. Calvin University 5 , Department of Mathematics and Statistics , , 3201 Burton St SE, Grand Rapids, MI 49546 , USA