1. Eye Center of Xiangya Hospital, Central South University, 410008 Changsha, Hunan, People's Republic of China
2. Department of Ophthalmology, The University of Pittsburgh School of Medicine, Pittsburgh, PA 15213, USA
3. Hunan Key Laboratory of Ophthalmology, 410008 Changsha, Hunan, People's Republic of China
4. Department of Bioengineering, University of Pittsburgh Swanson School of Engineering, Pittsburgh, Pennsylvania, United States of America
5. Department of Developmental Biology, Louis J. Fox Center for Vision Restoration, The University of Pittsburgh School of Medicine, Pittsburgh, PA 15213, USA