1. Vanderbilt University and Medical Center 1 Department of Biological Sciences , , Nashville, TN 37235 , USA
2. Vanderbilt University and Medical Center 2 Department of Cell and Developmental Biology , , Nashville, TN 37235 , USA
3. Vanderbilt Brain Institute, Vanderbilt University and Medical Center 3 , Nashville, TN 37235 , USA
4. Kennedy Center for Research on Human Development, Vanderbilt University and Medical Center 4 , Nashville, TN 37235 , USA