1. Hubrecht Institute-KNAW and Utrecht University Medical Center 1 , Utrecht 3584 CT , The Netherlands
2. Department of Pediatric Cardiology, Division of Pediatrics, University Medical Center Utrecht 2 , Utrecht 3584 EA , The Netherlands
3. Department of Biology, University of Padova 3 , Padova 35131 , Italy
4. Veneto Institute of Molecular Medicine 4 , Padova 35129 , Italy
5. Department of Anatomy and Embryology, Leiden University Medical Center 2333 5 , ZC, Leiden , The Netherlands
6. The Company of Biologists 6 , Bidder Building, Station Road, Histon, Cambridge CB24 9LF , UK
7. Division of Cardiology, Department of Medicine, Duke University Medical Center 7 , Durham, NC 27710 , USA
8. Department of Pathology, Duke University 8 , Durham, NC 27710 , USA