1. Department of Integrative Biology, University of Guelph, Guelph, Ontario N1G 2W1, Canada
2. Department of Biological Sciences, University of Alberta, Edmonton, Alberta T6G 2M9, Canada
3. Department of Biology, Acadia University, Wolfville, Nova Scotia B4P 2R6, Canada
4. Brevard County Environmentally Endangered Lands Program, Sarasota, Florida 34240, USA
5. Department of Biology, University of Ottawa, Ottawa, Ontario K1N 6N5, Canada