1. Virginia Commonwealth University 1 Integrative Life Sciences Doctoral Program , , Richmond, VA 2328 , USA
2. University of Richmond 2 Department of Biology , , Richmond, VA 23173 , USA
3. Nova Southeastern University 3 Department of Biological Sciences , , Fort Lauderdale, FL 33328 , USA
4. Center for Environmental Studies, Virginia Commonwealth University 4 , Richmond, VA 23284 , USA