1. University of Coimbra, Centre for Functional Ecology 1 , Associate Laboratory TERRA, Department of Life Sciences, 3000-456 Coimbra , Portugal
2. ECOMARE - Laboratory for Innovation and Sustainability of Marine Biological Resources, CESAM - Centre for Environmental and Marine Studies 2 , Department of Biology , , 3810-193 Aveiro , Portugal
3. University of Aveiro, Campus Universitário de Santiago 2 , Department of Biology , , 3810-193 Aveiro , Portugal