1. Department of Anatomy and Cell Biology, Graduate School of Medical Sciences, Kyushu University, 3-1-1 Maidashi, Higashi-ku, Fukuoka 812-8582, Japan
2. JST, PRESTO, 4-1-8 Honcho, Kawaguchi, Saitama 332-0012, Japan
3. Department of Advanced Information Technology, Faculty of Information Science and Electrical Engineering, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka 819-0385, Japan
4. Graduate School of Science, Kumamoto University, 2-39-1 Kurokami, Chuo-ku, Kumamoto 860-8555, Japan
5. Department of Radiology and Developmental Neuroscience Program, Saban Research Institute, Children's Hospital Los Angeles, Los Angeles, CA 90027, USA
6. Department of Radiology, Keck School of Medicine, University of Southern California, Los Angeles, CA 90089, USA