1. State Key Laboratory of Cell Biology, CAS Center for Excellence in Molecular Cell Science, Institute of Biochemistry and Cell Biology, University of Chinese Academy of Sciences, Chinese Academy of Sciences 1 , Shanghai 200031 , China
2. CAS-MPG Partner Institute for Computational Biology, Shanghai Institute of Nutrition and Health, Shanghai Institutes for Biological Sciences, University of Chinese Academy of Sciences, CAS 3 Key Laboratory of Computational Biology , , Shanghai 200031 , China
3. CAS Key Laboratory of Regenerative Biology, Guangdong Provincial Key Laboratory of Stem Cell and Regenerative Medicine, Guangdong Institutes of Biomedicine and Health, Chinese Academy of Sciences 4 , Guangzhou 510530 , China
4. School of Life Science, Hangzhou Institute for Advanced Study, University of Chinese Academy of Sciences, Chinese Academy of Sciences 2 , 310024 Hangzhou , China
5. Institute for Stem Cell and Regeneration, Chinese Academy of Sciences 5 , Beijing 100101 , China