1. Department of Genetics, Cell Biology and Development, University of Minnesota,321 Church Street SE, Minneapolis, MN 55455, USA
2. Developmental Biology Center, University of Minnesota, 321 Church Street SE,Minneapolis, MN 55455, USA
3. Department of Biochemistry, University of Wisconsin, 433 Babcock Drive,Madison, WI 53706-1544, USA
4. Howard Hughes Medical Institute, University of Wisconsin, 433 Babcock Drive,Madison, WI 53706-1544, USA