1. Osaka University Graduate School of Medicine 1 Department of Clinical Laboratory and Biomedical Sciences, Division of Health Sciences , , Suita, Osaka , Japan
2. Feinberg School of Medicine, Northwestern University 2 Department of Otolaryngology - Head and Neck Surgery , , Chicago, IL 60611, USA
3. The Hugh Knowles Center for Clinical and Basic Science in Hearing and Its Disorders, Northwestern University 3 , Evanston, IL 60208 , USA