1. Guangdong Provincial Key Laboratory of Insect Development Biology and Applied Technology, Institute of Insect Science and Technology, School of Life Sciences, South China Normal University 1 , Guangzhou 510631 , China
2. State Key Laboratory of Crop Stress Biology for Arid Areas and Key Laboratory of Integrated Pest Management on the Loess Plateau of Ministry of Agriculture, Northwest A&F University 2 , Yangling 712100 , China
3. Key Lab of Integrated Crop Pest Management of Shandong Province, College of Plant Health and Medicine, Qingdao Agricultural University 3 , Qingdao 266109 , China
4. Institute of Entomology, Guizhou University 4 , Guiyang 550025 , China