1. Kyoto University 1 Division of Applied Life Sciences, Graduate School of Agriculture , , Sakyo, Kyoto 606-8502 , Japan
2. Kyoto Pharmaceutical University 2 Department of Biophysical Chemistry , , Yamashina, Kyoto 607-8414 , Japan
3. Graduate School of Pharmaceutical Sciences, The University of Tokyo 4 , Tokyo 113-0033 , Japan
4. Institute for Integrated Cell-Material Sciences (iCeMS), Kyoto University 3 , Sakyo, Kyoto 606-8507 , Japan