1. Department of Molecular Biology, Yokohama City University Graduate School of Medical Science, Yokohama 236-0004, Japan.
2. Kihara Memorial Yokohama Foundation for the Advancement of Life Sciences,Yokohama 244-0813, Japan.
3. Department of Cell Biology, JFCR-Cancer Institute, Tokyo 135-8550,Japan.
4. Center for Translational and Advanced Animal Research (CTAAR), Tohoku University School of Medicine, Sendai 980-8575, Japan.
5. CREST, Japan Science and Technology Corporation (JST), Saitama 332-0012,Japan.