1. Department of Cell and Developmental Biology, Vanderbilt University, Nashville, TN 37232, USA
2. Vanderbilt Center for Stem Cell Biology, Vanderbilt University, Nashville, TN 37232, USA
3. Vanderbilt Brain Institute, Vanderbilt University, Nashville, TN 37232, USA
4. Vanderbilt Ingram Cancer Center, Vanderbilt University, Nashville, TN 37232, USA