1. Centre for Craniofacial Biology and Regenerative Biology, King's College London 1 , London SE1 9RT , UK
2. The Francis Crick Institute 2 , London NW1 1AT , UK
3. Randall Centre for Cell and Molecular Biophysics, King's College London 3 , London SE1 1UL , UK
4. Institute of Neurology, University College London 4 , London WC1N 3BG , UK