1. University of Cambridge 1 The Sainsbury Laboratory , , Cambridge, CB2 1LR , UK
2. Institute of Biotechnology, HiLIFE/Organismal and Evolutionary Biology Research Programme 2 , Faculty of Biological and Environmental Sciences , , 00790 Helsinki , Finland
3. Viikki Plant Science Centre, University of Helsinki 2 , Faculty of Biological and Environmental Sciences , , 00790 Helsinki , Finland