1. , Macquarie University 1 School of Natural Sciences , North Ryde, NSW 2109 , Australia
2. Ku-ring-gai Council 2 , Gordon, NSW 2072 , Australia
3. , University of Bristol 3 School of Biological Sciences , Bristol, BS8 1QU , UK
4. The University of Lincoln 4 , Lincoln, LN6 7TS , UK
5. Lancaster Environment Centre, Lancaster University 5 , Lancaster, LA1 4YQ , UK