1. School of Life Science, Arizona State University 1 , Tempe, AZ 85287-4501 , USA
2. University of British Columbia 2 Department of Zoology , , Vancouver, BC , Canada , V6T 1Z4
3. Virginia Tech 3 Department of Biomedical Engineering and Mechanics , , Blacksburg, VA 24061 , USA
4. University of Chicago 4 Department of Organismal Biology and Anatomy , , Chicago, IL 60637 , USA