1. Department of Ecology and Evolutionary Biology and Institute for Behavioral Genetics, University of Colorado at Boulder, Campus Box 427,Boulder, CO, 80309-0427, USA
2. Department of Mechanical Engineering, University of Colorado at Boulder,Campus Box 427, Boulder, CO, 80309-0427, USA
3. Department of Environmental Biology, University of Guelph, Guelph, ON, N0B 2J0, Canada