1. , University of Exeter 1 College of Life and Environmental Sciences, Biosciences , Exeter EX4 4QD , UK
2. Institute of Neuroanatomy, Mannheim Centre for Translational Neuroscience 2 , Medical Faculty Mannheim , , 68167 Mannheim , Germany
3. University of Heidelberg 2 , Medical Faculty Mannheim , , 68167 Mannheim , Germany
4. Mannheim Center for Translational Neurosciences 3 Division of Neurodegeneration, Department of Neurology , , Medical Faculty Mannheim , , 68167 Mannheim , Germany
5. Heidelberg University 3 Division of Neurodegeneration, Department of Neurology , , Medical Faculty Mannheim , , 68167 Mannheim , Germany
6. Umeå University 4 Department of Clinical Science, Neurosciences , , Umeå SE-90185 , Sweden