1. Teikyo University 1 Department of Biosciences , , 1-1, Toyosatodai, Utsunomiya, Tochigi , 320-8551, Japan
2. Institute of Life Science, Kurume University 2 Department of Cell Biology , , 67, Asahi-machi, Kurume, Fukuoka , 830-0011, Japan
3. Graduate School of Science and Engineering, Teikyo University 3 , 1-1, Toyosatodai, Utsunomiya, Tochigi , 320-8551, Japan