1. University of Turku 1 Department of Biology , , FI-20014 Turku , Finland
2. Université de Strasbourg, Centre National de la Recherche Scientifique, Institut Pluridisciplinaire Hubert Curien, UMR 7178 2 , 67087 Strasbourg , France
3. Institute for Marine and Antarctic Studies, University of Tasmania 3 , Battery Point, TAS 7004 , Australia
4. University of Jyväskylä 4 Department of Biological and Environmental Sciences , , FI-40014 Jyväskylä , Finland
5. Université Claude Bernard Lyon 1, CNRS, ENTPE, UMR 5023 LEHNA 5 , F-69622 Villeurbanne , France