1. Department of Biology, San Diego State University, San Diego, CA 92182, USA.
2. The Hospital for Sick Children, Program in Developmental and Stem Cell Biology, Toronto, ON M5G 0A4, Canada.
3. University of Toronto, Department of Molecular Genetics, Toronto, ON M5S 1A8, Canada.
4. Ontario Institute for Cancer Research, Toronto, ON M5G 1L7, Canada.