1. Microbial Genetics Laboratory, Genetic Strains Research Center, National Institute of Genetics, 1111 Yata, Mishima, Shizuoka, 411-8540, Japan
2. Yeast Genetic Resources Laboratory, Graduate School of Engineering, Osaka University, 1-6 Yamadaoka, Suita, Osaka, 565-0871, Japan
3. Department of Biology, Graduate School of Science, Osaka City University, 3-3-138 Sugimoto, Sumiyoshi-ku, Osaka, 558-8585, Japan