1. Faculty of Life and Environmental Sciences, University of Tsukuba, Tennodai 1-1-1, Tsukuba, Ibaraki 305-8572, Japan
2. Noto Marine Laboratory, Institute of Nature and Environmental Technology, Kanazawa University, Ogi, Noto-cho, Ishikawa 927-0553, Japan
3. Graduate School of Natural Science and Technology, Kanazawa University, Kakuma, Kanazawa, Ishikawa 920-1192, Japan
4. Research Center for Marine Biology, Tohoku University, Sakamoto 9, Asamushi, Aomori 039-3501, Japan
5. Department of Biology, Research and Education Center for Natural Sciences, Keio University, Hiyoshi, Kouhoku-ku, Yokohama, Kanagawa 223-8521, Japan
6. Misaki Marine Biological Station, Graduate School of Science, The University of Tokyo, 1024 Koajiro, Misaki, Miura, Kanagawa 238-0225, Japan