1. School of Biomedical Sciences, University of Liverpool, Ashton Street, Liverpool, L69 3GE, UK
2. Laboratory for Functional Morphology, University of Antwerp, Campus Drie Eiken, Universiteitsplein 1, Antwerp B-2610, Belgium
3. Centre for Research and Conservation, Royal Zoological Society of Antwerp, Antwerp B-2018, Belgium
4. Department of Anatomy, Medical School, Ruhr University, Bochum 44791, Germany
5. Faculty of Medicine, Katholieke Universiteit Leuven Campus Kortrijk, 8500, Belgium