1. University of Nottingham, Nottingham, NG7 2RD, UK
2. Center for Regenerative and Developmental Biology, and Department of Biology, Tufts University, Medford, MA 02155, USA
3. Micropathology Ltd, Warwick Science Park, Tile Hill, CV4 7EZ, UK
4. Wellcome Sanger Institute, Wellcome Genome Campus, Hinxton, Cambridge, CB10 1SA, UK
5. Community and Ecosystem Ecology, Division of Ecology and Evolutionary Biology, Graduate School of Life Sciences, Tohoku University, Aobayama, Sendai 980-8578, Japan
6. Department of Geobiology, University of Göttingen, Göttingen 37077, Germany