1. Botanical Institute, University of Cologne, Gyrhofstrasse 15, 50931 Cologne,Germany
2. Department of Molecular, Cellular, and Developmental Biology, University of Michigan, Ann Arbor, MI 48109, USA
3. Department of Botany, UF Genetics Institute, University of Florida, 220 Bartram Hall, PO Box 118526, Gainesville, FL 32611-8526, USA