1. Brown University 1 Department of Ecology, Evolution and Organismal Biology , , Providence, RI 02912, USA
2. 3D Anatomy Studios 2 , Providence, RI 02860, USA
3. Institute of Life Course and Medical Sciences 3 Department of Musculoskeletal and Ageing Science , , , Liverpool L7 8TX , UK
4. University of Liverpool 3 Department of Musculoskeletal and Ageing Science , , , Liverpool L7 8TX , UK