1. University of Michigan 1 Department of Earth and Environmental Science , , Ann Arbor, MI 48109 , USA
2. Texas A&M University 2 Department of Marine Biology , , Galveston Campus, Galveston, TX 77554 , USA
3. National Museum of Natural History 3 Department of Paleobiology , , Washington, DC 20560 , USA
4. Texas A&M University 4 Department of Ecology and Conservation Biology , , College Station, TX 77843 , USA