1. Department of Genetics, Evolution and Environment, and Institute of Healthy Ageing, Darwin Building, University College London, Gower Street, London WC1E 6BT, UK
2. Present address: Instituto de Bioquímica Vegetal y Fotosíntesis, Universidad de Sevilla-C.S.I.C., 41092 Sevilla, Spain
3. Spanish National Cancer Research Centre (CNIO), 28029 Madrid, Spain