1. Department of Biology, Faculty of Science, Yamaguchi University, Yamaguchi 753-8512, Japan
2. Present address: Division of Cell Biology, Institute of Life Science, Kurume University, Hyakunen-kohen 1-1, Kurume, Fukuoka 839-0864, Japan.
3. Department of Life Science, College of Science, Rikkyo University, Tokyo 171-8501, Japan
4. Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency, Gobancho, Chiyoda-ku, Tokyo 102-0076, Japan
5. Laboratory of Medical Mycology, Research Institute for Disease Mechanism and Control, Nagoya University School of Medicine, Nagoya 466-8550, Japan
6. Present address: Department of Microbiology, Aichi Gakuin University School of Dentistry, Kusumoto-cho, Chikusa-ku, Nagoya 464-8650, Japan.