1. Behavioural Biology, Department of Agriculture, Tokyo University of Agriculture and Technology, Fuchu, Tokyo 183-8509, Japan
2. Graduate School of Integrated Science, Yokohama City University, 22-2 Seto,Kanazawa-ku, Yokohama 236-0027, Japan
3. Laboratory of Neuroethology, The Graduate University for Advanced Studies(Sokendai), Shonan Village, Hayama 240-0193, Japan