1. F.M. Kirby Center for Molecular Ophthalmology, Scheie Eye Institute, Perelman School of Medicine at the University of Pennsylvania 1 , Philadelphia, PA 19104 , USA
2. Lewis-Sigler Institute for Integrative Genomics, Princeton University 2 , Princeton, NJ 08544 , USA
3. Princeton University 3 Department of Molecular Biology , , Princeton, NJ 08544 , USA
4. School of Dental Medicine, University of Pennsylvania 4 Department of Basic and Translational Sciences , , Philadelphia, PA 19104 , USA
5. CHOP-PENN Proteomics Core Facility, The Children's Hospital of Philadelphia Research Institute 5 , Philadelphia, PA 19104 , USA
6. Weill Cornell Medicine 6 Department of Ophthalmology , , New York, NY 10065 , USA
7. Weill Cornell Medicine 7 Department of Cell and Developmental Biology , , New York, NY 10065 , USA