1. Section of Molecular Craniofacial Embryology, Graduate School of Medical and Dental Sciences, Tokyo Medical and Dental University (TMDU), 1-5-45 Yushima, Bunkyo-ku, Tokyo 113-8549, Japan
2. Department of Physiological Chemistry and Metabolism, Graduate School of Medicine, The University of Tokyo, 7-3-1, Hongo, Bunkyo-ku, Tokyo 113-0033, Japan
3. Core Research for Evolutional Science and Technology (CREST), Japan Science and Technology Agency (JST), Chiyoda-ku, Tokyo, 102-0076, Japan
4. Friedrich Miescher Institute for Biomedical Research, Affiliated to the Novartis Institutes for Biomedical Research, Maulbeerstrasse 66, CH-4058 Basel, Switzerland
5. University of Basel, Petersplatz 10, 4003 Basel, Switzerland
6. Institute for Biology and Mathematics of Dynamical Cell Processes (iBMath), The University of Tokyo, 3-8-1 Komaba, Meguro-ku, Tokyo 153-8914, Japan
7. Evolutionary Morphology Laboratory, RIKEN, 2-2-3 Minatojimaminami-machi, Chuo-ku, Kobe, 650-0047, Japan