1. Institute of Biotechnology, HiLIFE, University of Helsinki 1 , Helsinki FI-00014 , Finland
2. Institute of Biomedicine, Cancer Laboratory FICAN west, University of Turku 2 , Turku FI-20014 , Finland
3. Turku Bioscience Centre, University of Turku and Åbo Akademi University 3 , Turku FI-20014 , Finland
4. Karolinska Institutet 4 Department of Cell and Molecular Biology (CMB) , , Stockholm SE-171 77 , Sweden
5. Finnish Cancer Institute, FICAN South Helsinki University Hospital & Translational Cancer Medicine, Medical Faculty, University of Helsinki 5 , Helsinki FI-00014 , Finland
6. University of Helsinki 6 Faculty of Biological and Environmental Sciences , , Helsinki FI-00014 , Finland