1. University of Alberta 1 Department of Oncology , , Edmonton, Alberta T6G 2E1 , Canada
2. Centre for Cooperative Research in Biosciences (CIC bioGUNE), Basque Research and Technology Alliance (BRTA), Bizkaia Technology Park 2 , Building 801A, 48160 Derio , Spain
3. Imperial College London 3 Department of Surgery and Cancer , , London W12 0NN , UK