1. School of Life Sciences, Peking University 1 Ministry of Education Key Laboratory of Cell Proliferation and Differentiation , , Beijing 100871 , China
2. Peking-Tsinghua Joint Center for Life Sciences, Academy for Advanced Interdisciplinary Studies, Peking University 2 , Beijing 100871 , China
3. China Agricultural University 3 Department of Entomology , , Beijing 100193 , China