1. Laboratory of Cell and Developmental Biology, Graduate School of Bioscience, Tokyo Institute of Technology 1 , Yokohama 226-8501 , Japan
2. Graduate School of Science, Kyoto University 2 Department of Zoology , , Kyoto 606-8502 , Japan
3. Faculty of Science and Technology, Kochi University 3 Department of Chemistry and Biotechnology , , Kochi 780-8520 , Japan