1. Sloan Kettering Institute, Memorial Sloan Kettering Cancer Center 1 Developmental Biology Program , , New York, NY 10065 , USA
2. Weill Cornell Graduate School of Medical Sciences, Cornell University 2 Biochemistry Cell and Molecular Biology Program , , New York, NY 10065 , USA
3. Wellcome Trust/Cancer Research UK Gurdon Institute, Cambridge University 3 , Cambridge CB2 1NQ , UK
4. Department of Physiology, Development and Neuroscience, University of Cambridge 4 , Cambridge, CB2 3DY , UK