1. Department of Otolaryngology – Head and Neck Surgery, Kyoto University Graduate School of Medicine, Kyoto 606-8507, Japan
2. Department of Speech and Hearing Sciences and Disorders, Faculty of Health and Medical Science, Kyoto University of Advanced Science, Kyoto 615-8577, Japan
3. Department of Biosystems Science, Institute for Frontier Life and Medical Sciences, Kyoto University, Kyoto 606-8507, Japan
4. Institute for Integrated Cell-Material Sciences (iCeMS), Kyoto University, Kyoto 606-8501, Japan
5. Department of Pathology and Biology of Diseases, Graduate School of Medicine, Kyoto University, Kyoto 606-8501, Japan
6. Research Center for Systemic Life Science, Graduate School of Biostudies, Kyoto University, Kyoto 606-8501, Japan
7. Department of Growth Regulation, Graduate School of Medicine, Kyoto University, Kyoto 606-8501, Japan
8. Department of Molecular and Cellular Biology, Graduate School of Biostudies, Kyoto University, Kyoto 606-8501, Japan