1. Broad Center for Regenerative Medicine and Stem Cell Research, University of Southern California Keck School of Medicine, Los Angeles, CA 90089, USA
2. Sir William Dunn School of Pathology, University of Oxford, Oxford OX1 3RE, UK
3. Department of Mouse Genetics and Metabolism, Institute for Genetics, Cologne Excellence Cluster on Cellular Stress Responses in Aging Associated Diseases (CECAD), Center of Molecular Medicine Cologne (CMMC), University of Cologne, Max Planck Institute for Neurological Research, D-50674 Cologne, Germany
4. Department for Internal Medicine, University Hospital Cologne, D-50924 Cologne, Germany