1. Nuclear Dynamics Group, Graduate School of Frontier Biosciences, Osaka University 1 , Suita 565-0871 , Japan
2. Laboratory of Chromosome Biology, Graduate School of Frontier Biosciences, Osaka University 3 , Suita 565-0871 , Japan
3. Laboratory of Mitochondrial Dynamics, Graduate School of Frontier Biosciences, Osaka University 2 , Suita 565-0871 , Japan