1. Memorial University, St. John's 1 Department of Biology , , Newfoundland and Labrador, A1B 3X9 , Canada
2. Centre for Ecology, Evolution, and Conservation, University of East Anglia 2 Department of Biological Sciences , , Norwich NR4 7TJ , United Kingdom
3. Soft Matter Lab, Memorial University, St. John's 3 Department of Physics and Physical Oceanography , , Newfoundland and Labrador, A1B 3X7 , Canada
4. ‡ Deceased