1. Biology Institute, Federal University of Bahia 1 , 40170-115 Salvador, BA , Brazil
2. Monitoring and Assessment Program of Natural and Artificial Ecosystems of Bahia State, Federal University of Bahia 2 , 40170-115, Salvador, BA , Brazil
3. College of Science 3 Department of Biology , , , PO Box 36, Al Khoud 123 , Sultanate of Oman
4. Sultan Qaboos University 3 Department of Biology , , , PO Box 36, Al Khoud 123 , Sultanate of Oman