1. Multicellular Organization Laboratory, National Institute of Genetics, Mishima, Japan
2. Department of Genetics, School of Life Science, Sokendai, Mishima, Japan
3. Department of Immunoregulation, Research Institute for Microbial Diseases, and WPI Immunology Frontier Research Center, Osaka University, Suita, Japan
4. Gene Network Laboratory, National Institute of Genetics, Mishima, Japan