1. Kavli Institute of Nanoscience, Delft University of Technology 1 Department of Bionanoscience , , Delft 2629 HZ , The Netherlands
2. Max Planck Institute of Molecular Physiology 2 Department of Mechanistic Cell Biology , , Dortmund 44227 , Germany
3. Utrecht University 3 Cell Biology, Neurobiology and Biophysics, Department of Biology, Faculty of Science , , Utrecht 3584 CH , The Netherlands
4. School of Biological and Behavioural Sciences, Queen Mary University of London 4 , London E1 4NS , UK